%PDF-1.4
%
1 0 obj
<< /S /URI /URI (http://www.pnas.org/lookup/suppl/doi:10.1073/pnas.1319538111/-/DCSupplemental/pnas.201319538SI.pdf?targetid=nameddest=ST1)
>>
endobj
2 0 obj 3535
endobj
3 0 obj
<< /S /URI /URI (http://www.pnas.org/lookup/suppl/doi:10.1073/pnas.1319538111/-/DCSupplemental/pnas.201319538SI.pdf?targetid=nameddest=STXT)
>>
endobj
4 0 obj
<< /F1 146 0 R /F2 203 0 R /F3 69 0 R /F4 136 0 R /F5 193 0 R /F6 58 0 R /F7 125 0 R /F8 184 0 R /F10 46 0 R /F9 115 0 R
>>
endobj
5 0 obj
<< /F18 117 0 R /F2 203 0 R /F3 69 0 R /F6 58 0 R /F8 184 0 R /F10 46 0 R /F9 115 0 R /F14 205 0 R /F16 139 0 R /F17 47 0 R
>>
endobj
6 0 obj
<< /Rect
[ 447.816986 344.750000 475.143005 351.722992
] /Subtype /Link /A 24 0 R /Border
[ 0 0 0
] /Type /Annot
>>
endobj
7 0 obj
<< /Subtype /Type1 /FontDescriptor 49 0 R /BaseFont /CLFEOM+AdvOT16d4f71d.I /Encoding /WinAnsiEncoding /Widths
[ 687 0 697 750 0 593 802 0 364 375 0 666 0 0 0 593 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 260 0 0 0 0 0 0 0 0 322
] /FirstChar 65 /Type /Font /LastChar 114
>>
endobj
8 0 obj
<< /Rect
[ 521.291016 495.382996 541.984009 502.355988
] /Subtype /Link /A 51 0 R /Border
[ 0 0 0
] /Type /Annot
>>
endobj
9 0 obj
<< /FontFile3 120 0 R /CharSet (/greater/less/equal/plusminus) /CapHeight 0 /Ascent 0 /Flags 32 /ItalicAngle 0 /Descent 0 /FontName /CLFEHL+AdvOTd877c31c /FontBBox
[ -62 -208 927 958
] /Type /FontDescriptor /StemV 0
>>
endobj
10 0 obj
<< /Subtype /XML /Type /Metadata /Length 199 0 R
>>
stream
Adobe Illustrator CS4
2014-07-08T20:12:42+05:30
256
148
JPEG
/9j/4AAQSkZJRgABAgEASABIAAD/7QAsUGhvdG9zaG9wIDMuMAA4QklNA+0AAAAAABAASAAAAAEA
AQBIAAAAAQAB/+4ADkFkb2JlAGTAAAAAAf/bAIQABgQEBAUEBgUFBgkGBQYJCwgGBggLDAoKCwoK
DBAMDAwMDAwQDA4PEA8ODBMTFBQTExwbGxscHx8fHx8fHx8fHwEHBwcNDA0YEBAYGhURFRofHx8f
Hx8fHx8fHx8fHx8fHx8fHx8fHx8fHx8fHx8fHx8fHx8fHx8fHx8fHx8fHx8f/8AAEQgAlAEAAwER
AAIRAQMRAf/EAaIAAAAHAQEBAQEAAAAAAAAAAAQFAwIGAQAHCAkKCwEAAgIDAQEBAQEAAAAAAAAA
AQACAwQFBgcICQoLEAACAQMDAgQCBgcDBAIGAnMBAgMRBAAFIRIxQVEGE2EicYEUMpGhBxWxQiPB
UtHhMxZi8CRygvElQzRTkqKyY3PCNUQnk6OzNhdUZHTD0uIIJoMJChgZhJRFRqS0VtNVKBry4/PE
1OT0ZXWFlaW1xdXl9WZ2hpamtsbW5vY3R1dnd4eXp7fH1+f3OEhYaHiImKi4yNjo+Ck5SVlpeYmZ
qbnJ2en5KjpKWmp6ipqqusra6voRAAICAQIDBQUEBQYECAMDbQEAAhEDBCESMUEFURNhIgZxgZEy
obHwFMHR4SNCFVJicvEzJDRDghaSUyWiY7LCB3PSNeJEgxdUkwgJChgZJjZFGidkdFU38qOzwygp
0+PzhJSktMTU5PRldYWVpbXF1eX1RlZmdoaWprbG1ub2R1dnd4eXp7fH1+f3OEhYaHiImKi4yNjo
+DlJWWl5iZmpucnZ6fkqOkpaanqKmqq6ytrq+v/aAAwDAQACEQMRAD8A9JaV5q0fVdQuLGy+tPLa
tLHLM9ldxWxe3l9GVUupIkt5CsgIojnoadDiqb4q7FUnk07T7zzBdfXLWK59O0teHrRrJxrJcVpy
BpWmKoDzhottbeUdbudG0q3fWIbC6k01Y7WKVzcpCxhCxlG5n1AKLQ1xV5rpeta7b3OiaadIj1C7
1GWd7uK+0qY3AhinsYX9Nxp+iLDHFFdyTGSW3dSy+mHLEAKofRPMn5htbW8OoeSrcXCxWZeV9FuK
yzSQgvbsyGNI3kcEtPwEMP2WBJ2VQkvmD8yV84Sr/heJNC+pn1T/AIduZYbS5E268UpPfOFpHzhk
WMg+oForYqyrztc+c9Pu2Hl3ytp1xbxWUM7JLpUtyz3D219cSRpJDNEo4vZQw8eJPKZTXopVSG18
y/mTqN8be28h2umxLPwkuNQ0WQqoe7trf009K6/eCGO4eZp6qsioeKrRiFUxs9Y883hghPk2xspr
iDT+by6TO8dvJdNYLPI59SISKv1q6rEGVohBV2IYYqy38vKa/pV3d6xoWn2kltdyWCJFbKoeSzCw
3cg5GT4DdpMqCuyqKkmuKpj5s0DQU0uApptqpOo6YKiGMbNqEAI+z3Bpiqc/4d8v/wDVstP+REf/
ADTirv8ADvl//q2Wn/IiP/mnFWK+f9J1G1h0VvK2jWk9w2oj9IQG1iZJLSO1uJniZyh9H1ZI441k
24sy9tiqwrS9X8+PZ2FjN5SthqMtrYvLfXOjzMge4/RkcskqxG3i5Rvc3peIMjKIgSAN2VS+885f
mRbW0Sr+XEVzeNbmZfR0eX0XkksorkRuzThoDbySmNtnMzKVXg2wVTltc83RT6xZ3HlO0U2D2iW+
pR6FeTQsGEouJEgjd3uQXjj4CKSqLJ8VeDYqyn8tnPmLRbi+1vy/YWE8Ny9kIEtkFZbRVhu2qWlH
H64syoAx+FRUk4qy3/Dvl/8A6tlp/wAiI/8AmnFXf4d8v/8AVstP+REf/NOKpN5T0DQX0ucvptqx
Go6mKmGM7LqE4A+z2Apiqn510e3tfLs02kaXCLoS2yyyW9nFcTx2z3Ea3UsMJjl9SSO3Lui8GqR9
lvslV5v5g1bznaqdO0Hy3LfQy6Zez22tXuhcLr14ku5ICyRKkCu31aONYpIEZjIrcP2Cqg9R/MDz
dZ6XqNxdeS7XTp7Rp2tmu9FmZZjClfq8cUNw8jJWn+mnjFuBwFcVTRPMnm+LS21bUPKWn2djp0E+
oasJNKmjZoLVbKSa3hMsqCOVBc3SJIwZZfR5qoU4qq+b38/WFz5VutN8u2s4n0nUbrXdOttME0Iv
4rIzW0EklHlUGekaqrKzGvWoCqoGXzf55utaex0zyAkdrNdGGz1C70SdYFtXv/QjvJWeaCUUtyC0
BiVusnIJtiq/UvNXnTTiTP5OsCYp5LH0v0TOiTyPM9lZ3aXTyelHHPdtD/o9Hl9MlgStGxV7Cnlz
QgiiTTbNnAHJlt41BPcgENT78VUI9N06z8w2htLWG2L2l0HMUapWktvSvECuKvKfyh/wnd/mZrWs
6UsQvdSivZ5WR9OllKy3ccjCZrPU9Q+wxCoUt4lYbuXahxV7Lq2owaZpd5qU6s8FlDJcSpGAXKxK
XIUEqKmm1TiqT6L540zVtZXSIYJobw2C6i6zNbgopuHtmiKLK8jOkkTBmRWjG3x/EKqrWsfLGveY
bw3NvZaqba1tlBkSK49MtLcVHxBuNeOKor/BHkv/AKsGm/8ASJB/zRiq0+RfJBcOfL2mcwCob6nb
1ANCRXh3oMVXf4I8l/8AVg03/pEg/wCaMVd/gjyX/wBWDTf+kSD/AJoxV3+CPJf/AFYNN/6RIP8A
mjFXf4I8l/8AVg03/pEg/wCaMVd/gjyX/wBWDTf+kSD/AJoxVbH5E8jxIEj8u6YiL9lVs7cAfQEx
VKfNfk3yhHpcDR6Hp6MdQ01SVtYAeLahArDZOhUkHFU4/wAEeS/+rBpv/SJB/wA0Yq7/AAR5L/6s
Gm/9IkH/ADRirv8ABHkv/qwab/0iQf8ANGKu/wAEeS/+rBpv/SJB/wA0Yq7/AAR5L/6sGm/9IkH/
ADRirv8ABHkv/qwab/0iQf8ANGKrY/InkeJAkfl3TERfsqtnbgD6AmKrv8EeS/8Aqwab/wBIkH/N
GKu/wR5L/wCrBpv/AEiQf80Yqk/lTyb5Qk0udpND092GoakoLWsBPFdQnVRunQKABiqcf4I8l/8A
Vg03/pEg/wCaMVd/gjyX/wBWDTf+kSD/AJoxVTl8g+RJuHq+XNLk9Ng6c7K3biw6EVTY4quk8ieR
5UKSeXdMdDSqtZ25BoajYpiq7/BHkv8A6sGm/wDSJB/zRirv8EeS/wDqwab/ANIkH/NGKrX8ieR3
48/LumNwIZOVnbmjDoRVOuKrv8EeS/8Aqwab/wBIkH/NGKoP9H+VdA8wWk0FtY6T69pdI0iJDbc6
SW5CkgJWmKsP8g+VbKw/M3WdWi0q9tbu5/SYu7iePhAPVv0kjKzfU4PrBuEHqLxuJPTUcW+I8sVe
pzwQXEElvcRrNBMrRyxSKGR0YUZWU1BBBoQcVQdl5f0GxuVurLTbW1uUgW1WeGCONxbpTjCGVQfT
XiKL0xVqD/lIL3/mEtP+TlziqYYq7FXYq7FXYq7FUumuNcXXra3is4X0V7eV7q/MxEqTqyCONYeP
xBlLGvL7tuSqY4qkvm7/AI5UH/bS0v8A7qNviqdYq7FXYq7FUv1W41yG405dMs4ruCa5EepPLN6R
htijEyoOLc2DBRx9/pCqYYq7FXYq7FUl8o/8cqf/ALaWqf8AdRuMVTrFXYq7FXYq7FUukuddHmGG
2jsom0NrV5J9QM1JVuQ6hIhDx3UoS3Ll937SqY4q7FUvn/5SCy/5hLv/AJOW2KvJfyt0bVLf82de
utRvNLub2KK/ju0sLbS7SRTPeRSRsRZ3E17IHSOr/XEDI2wZySxVe1Yq7FUvg/5SC9/5hLT/AJOX
OKphirsVdirsVdirsVQ1tqWnXVxc21rdQz3FkwjvIY5Fd4XZeSrIoJKEqagHFUTiqS+bv+OVB/20
tL/7qNviqdYq7FXYq7FVK6ura0tpbq6lSC2gRpJ55GCIiIKszMaAAAVJOKrre4guYI7i3kSa3mRZ
IZo2DI6MKqysKggg1BGKr8VdirsVSXyj/wAcqf8A7aWqf91G4xVOsVdirsVdirsVQ9vqWn3N1c2l
vdRTXVkyreQRurSRM68kEig1Xku4riqIxV2KpfP/AMpBZf8AMJd/8nLbFWDfl7e+WLzzdqcsGgza
R5lWOYaokmp2l4sJe45yRrbQXtw8PqSsZCRbopNTWp3VZ3rumfpXRNQ0z1PR+vW0tt6vHnx9VCnL
jVeVK9KiuKsa8t/l3Jo3mW31ttQSZbbSf0SlnFbmGNVNx9Y5J+9k4opJVI6fCtByNN1U8063mh8w
ah6l1Lc8rW0I9URjj+8udh6aR/jXFU3xV2KuxV2KuxV2KpPoOgSaZeavdzXEd1Nqt2bkOttFbukQ
RUjhd4wDN6YU0d96bYqnGKpL5u/45UH/AG0tL/7qNviqdYq7FXYq7FVO5tre5t5ba5iSe3mUxzQy
KHR0YUZWVqggjYg4qhL/AE2ZtDn03SLgaRL9XMFhcwxI62xCcY2SE0QhNqL07YqibKK4hs4Ibmc3
VxHGiTXJVUMrqoDSFF+FeR3oNhiqtirsVSXyj/xyp/8Atpap/wB1G4xVOsVdirsVdirsVQ8Gnafb
3NzdW9tFDdXhVrueNFWSYxrxQyMBV+K7CvQYqh3sNVPmCO/XUiulLatC+keihDTlwy3HrfbFFBXh
074qmGKpRqVvNPr1gIrqW1ItbslohESf3ltsfVSQfdirD/I17YXfnfVXsvL11aW0L6jCNXlubya1
M8d8FuUt4JV+qxevIPUYwtu6sGFVJxV6PirsVS+D/lIL3/mEtP8Ak5c4qmGKuxV2KuxV2Kqdzc29
rbS3NzIsNvAjSTTSEKiIg5MzMdgABUnFVLTNTsNU0+31HT50ubG7jWa2uIzVXRxUEYqicVSXzd/x
yoP+2lpf/dRt8VTrFXYq7FXYq7FULquq6dpOnXGpalcJa2FohluLiQ0VEXqT/Tviqtb3EFzbxXFv
Is1vMiyQyxkMjo4qrKw2IINQcVVMVdiqS+Uf+OVP/wBtLVP+6jcYqnWKuxV2KuxV2KoLWdb0jRNP
fUdWu4rGxjZEkuZmCorSOEQEnxZgMVRuKuxVL5/+Ugsv+YS7/wCTltiry78u7RE/N7zBMLrTpGZN
QYx2tgttOwa/Va/WBp9r6wjMZWU/Wpf3nXccsVek+a9X1bS9HnudK0qfVb0JJ6UcAiYRsEJV5UeW
B3TkBVYuTn9lcVSbyx5n80ah5hjsNR06SCxOlJdm8+o3FvG12bh0ZfUmk/dlofTkEBRnWrcnBWhV
TrTriabzBqHqWsttxtbQD1TGeX7y53HpvJ+NMVTfFXYq7FXYq7FULY6lpeqW8kljcw3turvBK8Lr
KgkjPGSNipIqp2IxVXhhhghjggjWKGJQkUSAKqqooqqo2AA6DFV+KpL5u/45UH/bS0v/ALqNviqd
Yq7FXYq7FVK6ura0tZrq6lSC2t0aWeeRgqIiDkzMx2AAFScVWq1jqNiGUxXdheRVBHGSKWGVfpVk
dT8iMVVwAAABQDoMVdirsVSXyj/xyp/+2lqn/dRuMVTrFXYq7FXYqhV1XTG1N9KW7hOpxxC4ksg6
+sIWYqJDHXlx5ClaYqrz28FxEYp40miJBMcihlJUhgaHbYioxVfirsVSjUriaDXrAxWst0Ta3YKx
GIEfvLbc+q8Y+7FWBeWvLljB+Yes3+h+a9I/StxcNJqeh20UFzdeityPXFzNJNJdht+IC8I42IHE
gAYq9UxV2KpfB/ykF7/zCWn/ACcucVTDFXYq7FUvs9ZS61jUNLFndQtpwhZruWIpbTeupalvLX95
wpR/A4q1pkXmBL/U21K4tprGSZW0mOCN0kjh9NQ6zMzMGb1K0oP10VVF2ljZWUbRWdvHbRO7yukK
LGpkkYs7kKB8TMak9ziqtirsVSXzd/xyoP8AtpaX/wB1G3xVOsVdiqC1vVF0nSbvUmtri8W0jaU2
tnH61xJx/ZijqOTH54qpaq2s3fl+dtCeO01aeDlYvfRuUjkZar60YIYU7+B7HpiqMNsLiyNtfpFc
CWL07qMp+6kDLxkHpuX+Bt/hJO2Kr4IIYIY4II1ihiUJFEgCqqqKKqqNgAOgxVfirsVdiqS+Uf8A
jlT/APbS1T/uo3GKp1iqndC5a2mFqyJdFGEDyKXRZKfCXUFSyg9QCPniqH0aPVo9Js01iWGfVVhQ
X01srJC0wHxmNWJIWv8AmMVU9f1hdG0i41NrS6vltwpNpYxGe4fkwX93GCOVOVTv0xVEpZWX1tr9
beNb2SNYXueCiYxKSyxs9OXEMxPGvXFVfFXYq7FUvn/5SCy/5hLv/k5bYq8q/LTWdQl/NrzDpXov
b6dCNTmjUS6gbd2OohOcS3F5Pauxf1C5ht04tVdjyXFXsNzc29rby3NzKsNvAjSTTSEKiIg5MzMd
gABUk4qltl5r0C91ODS7a7El9c2KapBDwkHKzkYIsvIqFFWP2SeXtiq+znhl8wX3pSLJxtLQNxIN
D6lz1piqZ4q7FXYq7FWMX91aax5L1Q+brOTy/pzrcwXyTXKIy2yOyCX14HookQBqV703HVVryzb6
Hb+YtWOladdRi6t7GaXWnd5bS8VYjHAIJHkk5FI1HMgb1qa7EqsoxV2KpL5u/wCOVB/20tL/AO6j
b4qnWKuxV2KoOPWdJk1WXSEvIW1SCJbiaxDqZlhclVkKV5cSR1/riqMxVLrDQNOsNU1LU7dZBd6s
0T3rPLI6EwR+mnBGYonw9eIFe+KpR5TWHT9b13QbLQ7nTtOtZkvE1OZ2khvJ74NNOYTIWb4G2YA0
B7DaqrKMVdiqS+Uf+OVP/wBtLVP+6jcYqnWKuxVCatplrqul3emXfP6rewvbziN2jfhIpVqOpDDY
4qq2dnBZ2UFnbhhb20aQxBmZ2CIoVauxLMaDqTXFULoOg6VoGkw6TpMJgsLbn6MJd5CObmRvikLN
uzE7nFUj0vz817rOkabcaRc6WdYtbq4hGoSQQXCyWkoRofqvNpWqn7zku3GnvRVlmKuxVK7yeCHX
7EyyLGDaXYBcha/vLbxxVgP5eW+qf8rE8w3l9YNayTyXy/WDY/VhLFHe8bcmb9E23qVhCkH9Iy8v
tcf5FXo2o6NpGp/V/wBJWNvem0lWe1NxEkpimX7MsZYHg69mXcYqg7Dyh5esNVj1W0tTHfw2SabF
KZZXC2kZDLEqO7IACo3pX3xVVs4IYvMF96Uax8rS0LcQBU+pc9aYqmeKuxVL78+YBqemjT1tTphe
T9LmcyeuE9M+l9XC/BX1Ptc+3TFWrS+1eXWdQtLjTfq+m2ywmx1H1kf6y0ikyj0l+KP0iAPi+12x
VrzLp97qPl/UbCxkgiu7q3khhkuoRcQBnUj95E3wuu+4NR7Hpiqnoi+ZknvItXFkbOIwrpktmJEd
1ES+s0sbllT97XgFY/DselSqm2KuxVJfN3/HKg/7aWl/91G3xVOsVUb7679SuPqHpfXvSf6r6/L0
vV4nh6nD4uHKnKm9MVSry3feapzcW/mDTIbOW2S3CXttMJILqR4g1wYoz+9iWOWqjnuRviqrDbas
PNVzdPa2K6W1nFFFeIG+vvMsjs0ch48fRUNVRXqT47Kqtxb682vWlxBeQx6JHBMt5YtCWmknYp6T
rNyHEKA23H76gqqmOKuxV2KuxVJfKP8Axyp/+2lqn/dRuMVTrFXYql2sX2sWr2A03Tf0itxdRw3z
eskH1a3YHncfGD6nCg+BdziqX6lB58uTqsNldafp6epanRLsxyXEnpgg3QuY2KJVqFU4HofHfFU3
1ZtUXS7ttJWF9UELmxS5LCEzcT6YkKfFx5daYq1bWhlSyutSgt21a3io00a8hHJIq+sIXccwjFfa
oAriqLxV2KpZeQQza/YrLGsgFpdkBwGFfUt/HFWH+RL380rjzZqR8z6Xe2OhuLl9Pe4u9JkhANwD
bRi3s4Tcq6wGhZ7l1qD1qKKvQ8VdiqXwf8pBe/8AMJaf8nLnFUwxV2KuxV2KpffXWtR6rpsFnYpc
adOZv0leNMI2twiVi4xcSZPUf4TQjjirXl+11u10uOHW76PUtRVpDJdxQi3VlaRmjHphnpxQhevb
FUxxV2KpL5u/45UH/bS0v/uo2+Kp1irsVWTrM0MiwOscxUiKRl5qrEfCSoK8gD2qMVQmhW+r22j2
cGsXiX+qRxKt5eRRCBJZB1YRgkL/AJ7DpiqzWLvWrc2P6LsEvhNdxxX/ADmEJgtWr6k61B5lNvg6
nFUxxV2KuxV2KpL5R/45U/8A20tU/wC6jcYqnWKuxV2KpbrF3rtvPpy6XYR3sM90sepPJMITBbFW
LTKCreoVYD4R1/EKt6ja65LqOmS2F9HbWEEkjapbPD6jXEbRlUVJOQ9Pi/xdN/wKqY4q7FXYql8/
/KQWX/MJd/8AJy2xVgPkeDzcv5la1PqNxqD6XIt8q2tyt8LeEper9VZHnP1OT1bckr9VA4KKPVqs
VWfa1pM+p26QR6ldacnKsxtDErSxkENEzvHI6Ka/aiZHH7LDFUj0HyNPpXmSDWG1CO4ig0W20Rbf
6qiyEW0hkExuCzyAMWasa0U7E1KjFU10+3lh8wah6lzJccrW0K+qIxxHqXOw9NI/xriqb4ql2pr5
hN7pp0trVbITH9LC5EhlMHA8fq5Q8efOn2tqfiqv1sa2dJuhoZthq/D/AEM3okNv6lf92el8fH5Y
qpazoFlrmmxWOrBnRZILhhBLLB+9t3WVSGjZX481rSuKpnirsVdirsVSXzd/xyoP+2lpf/dRt8VR
mujXDpFyNBNsNX4/6Gb31Pq/Oo/vPS+OlK9MVWeXx5gGlxjzA1q+q8pPWNgJBb8ebenxEpL/AN3x
rXviqY4qhtS0+21LTrrT7oM1reRPBOqMyMUkUqwDKQw2PUYq7TdPttN0610+1DLa2cSQQK7M7BI1
CqCzEsdh1OKonFXYq7FXYqkvlH/jlT/9tLVP+6jcYqnWKpfri6+1ko0J7VL31ouZvVkaL0PUHrUE
ZDc/Trx7V64ql3mnQL7XprPTZhA/lqTk+rJ6lxDdmSJ0ltfQeFlHESJ8fI9OmKshxV2KuxV2KuxV
KNRt5p9esFiupbUi1uyWiERJ/eW+x9VJBirzv8tLOBPzL8x3FtpVxHbM+of7mJIbVo5JW1AiWL61
Hp9tK55oSE+uTBV+F6OtFVet4q7FUvg/5SC9/wCYS0/5OXOKphirsVdirsVdirsVdirsVSXzd/xy
oP8AtpaX/wB1G3xVOsVdirsVdirsVdirsVdirsVSXyj/AMcqf/tpap/3UbjFU6xV2KuxV2KuxV2K
uxV2KpfP/wApBZf8wl3/AMnLbFXjH5Ja8+u/mNrOqCy0u0t762ubiP6rJokl5V7tW/eCxpfAcWXm
ZmcO45VWqrir13zdrGq6TotxdaZplxqV2I5PTW2jjm9JghKySQNNbSTICN44SZG6KMVSPyr5r836
nr9lZ6hpbW+my6Mt7c3jWd1alb5pgvofvyVQ+kQxj+Jga/EQMVTWXWobC91vU9RhksrPT9Pgnnkl
MRrHE107MvB37L3piqVflP5+m85+X57u8NmNTs7gw3ceny+tbgSIs8PF6tUiKVUff7at06YqzbFX
Yq7FXYq7FXYq7FUl83f8cqD/ALaWl/8AdRt8VYTdfmV5ih853WmL9X+oW+uWmjrbtp94OUVzFbu0
rar6v1JJVa4PGFk5NQKN3BxV6hirsVdirsVdirsVdirsVefJruu250/QdCFtHqWrahrs31y+SSW3
ihs9Qk5j0onhZ3dplCjmNqntQqsv8vXmo3WmK2pG1N/FJLBcmycyQFoZGSq8viQkKCyMSUPw1NK4
qmWKuxV2KuxV2KuxV2KpRqNxNBr1g0VrLdE2t2CsRiBH7y33PqvGMVYH+VzaguuIZo9S4XdjLPPF
eReYoY7KUyRN6DPq889tcyMZG/eRojfATSjGir1PFXYqkOu6FJqUt1FNaQXun3kNvHNBLcS25Jtp
nlAPpRvyUlxUVodwQQcVQ+j2k5mOvWWkWdvdapa26zst3MvKKPnJEGjEHp8l9dviAr2rQDFUfcaj
rsE1tE1jak3UpiQi7k2IjeWp/wBH8Iziqv6/mD/litP+kuT/ALJsVULfUddnluYlsbUG1kETk3cl
CTGktR/o/hIMVdfajrtnZXF3JY2rJbxvKyrdyVIRSxArb+2Kq/r+YP8AlitP+kuT/smxVQl1HXo7
mCA2NqWn58SLuSg4Cpr/AKPiqv6/mD/litP+kuT/ALJsVSnUf8Q6xZmCGztI2tr61kZnupaH6ndR
TsBS3/aEdBiqW/4StJvM9zetpMZvxLb6nMp1a/8Aqj3NDDFMbTh9WMiC1WjGPainqMVZR6/mD/li
tP8ApLk/7JsVULHUddvLK3u0sbVUuIklVWu5KgOoYA0t/fFXXGo67BLbRNY2pN1IYkIu5KAiN5an
/R/CM4qr+v5g/wCWK0/6S5P+ybFVC31HXZ5rmJbG1BtZRE5N3JuTGktR/o/hIMVVJ7zX4YZJWsbQ
iNS5Au5K0UV/5Z8VdDea/NDHKtjahZFDgG7krRhX/lnxVTl1HXo7mCA2NqWn58SLuSg4Cpr/AKPi
rHNQ8szyaMltrOm21xx1Gee2mttQuraaJtQvXK8JoYYpU+G4CvxbcV64qn2k6ff6Rp8Wn6bpdlbW
cFfTiW7lO7MWZiTblmZmJZmJqTucVVbPUdduomlSxtVCySxEG7krWGRoidrfuUxVz6jrq3sVobG1
5zRySq31uSgETIpB/wBH/wCLRirrzUddtbZ53sbVlSlQLuSu5A72/viqv6/mD/litP8ApLk/7JsV
ULbUdduPV42NqPRkaJq3cm5WlSP9H98VdLqOvR3MEBsbUtPz4kXclBwFTX/R8VVJ7zX4YZJWsbQi
NS5Au5K0UV/5Z8VWWS6pdX9rqFzBBBAttKirHM8rlp2icVDRRAACM98VTbFXYq7FUj82S+bVseHl
62ild1b15zOsVzEART6rFLDLbyyMOXH1pEQGlaiuKpf5QuvPcmpSw+YbJrawi06yWCblasJL1Q/1
x2EDllLMygLx4UWo60xVQ1qf8y/8Q6W1nZQposb363y2s8Nw8kfogWUkguo7Ro39QklImYDuxGKp
h5Wbze3kuNtYR4vMjR3B9K8a2dlkMjmBZHswsDhV4DkirUblVNRiqVeWG/Mc6lpY1aOVNPEN1+kW
uPqHqO/qn6uZPqpP7z0uAHpUTZ+W/DFU286HzcItIPltPUI1O3Orxj0Ktp9G9dR9YIXf4fsnl4Yq
7yTJ5qe11JvMcU8Mxv5fqEdwbRiLQKgjKmzPHgzcmAk+Na8SWoGKrH/MV1+bVveeaDo9j9cgY2A8
s72QCLxH14/vZIWLdaeqaV6VGxVZprv6T/Qmofor/jp/VpfqP2P7/gfT/vPg+1T7W3jtirF9Dm/M
oavS+tOWnx6QTEbma0iEup+uSq3Bto5nRvR4h3jUxhuXFT8OKoGaf83vXvCLIBpLux9NYZ7NoIrc
TN9aFvLJHHNIvpFOfrRqxPqenx+DFWU+a5fNi6bcJ5ctreW6e1ueE8s/pyxXAiP1b0omhkil5Sdf
UkQD3xVKfK0nn86zZLrlvJHpo0O2F05ez9MasspFx+7hVpSXQqVKuIwB9mp+FVvWrn8x184aU1jY
QyeV4frP130LiJ5pqwAQeqlxFCYeMvT0ZHqK8qbAqo7ypcedH8m28+uWkS+aikpms5ZEhh9T1X9N
TJbfW1RCnHdeZA61NcVY9C/5sHh60LxTSa1A8oiewkhTTxMfrCCRhFI0HoceDGMTlwahRTFWQedp
POY0/wBPy1bJIzlfrFxHPEl5GokXkLaG5he1kZk5D97KgHudsVW+V384NrOr/pyOSOwWOzXTwfqp
gMgiIumg9EmcKZKbT7/y7bBVC3Mv5jv5ijK2ttb6UIbmNDDKl3HzNBBNOJUspw9SD6cTFeIerlim
KoW8m/NHjoqrZxelE1m2pS209u9xM1IzdCeG4iiiijH7wAwTM5bgRxXliqL0y9/MiTyu8s+mWyeY
nurkfVry5WKKK3aSRrdke1juxLwQxrRuBbcsVOKoXy035jfpDSF1SOZbAQ336Ua5NgHL+tW0Li1r
+8MbdYqIOLcgSVIVQOtXH5wkatJBYCOQ29uNMg0u5spgsvrAyFJ9QhtzVow3q+pFxpw9P4uRxVOP
ND/mJ9UsotDtreT/AHma9u2mjW558/3qfVZIng9LoXYT8+PIIOXE4qjWPm86N5gbYakzTtoKqsSl
UNuhgVgXlQus3IMXahO+y0GKpZ5aP5iHzLbjV0ZNBXSqTGX6n6jah9YqrP8AVyW9T6uQH4fuuQPH
tirepf8AKxf0n5gGnj/RjCg0N5VtXiWQRITwAeObkziRWM1VBKFQVDjFUx86N5uWPSG8tJ6hGp25
1eMehVtPAb1wPXKjf4fsnl4Yq15JfzU9rqTeYopoZfr8osI5zaMRaBUEZQ2Z48GbkwD/ABrXiS1A
xVZHirsVdirsVdirsVdirsVdirsVSvzLr8GgaQ+pT2812qy29vHbW3p+rJLd3EdtEq+q8UYrJMtS
zgAYqw7Wfz08p6RbmS6stSaZFkNxawQJNJC8JvVmjkKSGMNG+lTKxDlejBinJlVTa2/NjyLcW8M8
d7MROkkioLO7ZgsUcMpLBImpzW6h9I9JfUT0+XJaqtw/mx5EmadY76YvbheamyvQS7NCghQGEepP
zu4VMKVkBYArXFVeP8yfKU+javq1pcyXNtols15fIsMsUnpKJN0E6xBvit5E2NA6spoykBVrT/zN
8lahpVxqlpfSSWlsLcvW0u0lcXj+namGB4lmmE8nwxmNG5HYVOKoa5/N78vraN5JtRlVUUuB9TvS
zqsbSymNRCWk9FEYzcAfSofU44qo6r+cfkrTpY4i93cyvqUOkFIbSdSLieWaFWX1liE0YktJVLQ8
91IFcVX2P5w+QLvRoNXGoPFbzwpOY3gmMsaSev8A3ixo9OIs5nY/ZCIz14fFiqYX35ieULDU73TL
q8kjvNPoLlfq1yyc2iWZYY5FiMcszRyKyxRsznsuKpRJ+dfkJdUh06Ke7uJZAfWeKyumWFqRcY5A
Yw5lZ7mKMRIrOHdVZQTiqvP+cf5eQyiP9ITy8lidJLewv542WdYWj4yQwOjFvrkIoDWsir1YDFXS
/nL+XMc9vb/pOSW4u4oZ7SCGzvZpJo7iEXCNEkULs/7ohm4j4K/FQ4qrt+a/kEWz3f6TLWi8eN0l
vcvDIGjaYtFKsRSVY443eUoSIwpL8aHFWW4q7FXYq7FXYq7FXYq7FXYq7FXYq7FUl85ebdL8o+XL
rzBqqytYWZiEwgUPJ+/mSBSFJWtGkBO/TFUot/za8kTajNYfXGjmj1KTR4XMbNFNdxJEzqkkfNFH
K4SMGQrV9hXFURoX5meUdb0x9RtLiVIYrFdTnWWCZStuYxK/FgrRzNEGCyLCz8W+E74qgT+c35ft
c6VBBezXC6vcC0guY7W49CKZg5CXErRqsT1joyN8a15MoSrBV3/K6vy29aKBNTllnuFR7WGKxvpJ
J0kjMqtAiQM0q8BVmQEL3pirtP8Azn/Ly8RXGoPBHILt4JpYJhDJFYmb1ZUnVGhKFbWR0+PkVHSu
2KqzeZvIPnWI6A01xOLh+ZiMV/YOk9jPz4esUt2huIpbfn6fNZKLypx3xVKLnT/yUuNDm1BZINV0
6HhYTzWFxc6lK0l2biBUf6o880k0zarPVjV2MhYmtCFUngsP+ceLLTJtKtrQGzv7aSRrOGLUpnli
vxaXMqxqivI0gRLWRkT44l4sQgOKoL9Jf8496RowXSrGeWziX1lWwW+jkjE1ouoJcJI7wsC6aJGw
lRuSyJUlTzbFU70G3/Iyytb7TtJiWOHzb6VpqMYF8xnM8UXBJXephqNRiViStHlVW+NgMVRF/f8A
5M6Vp9xYXMptbW3jstMuOH18PAmm3NytlWWP44fRntp2SbkPs8+XHicVYprmlflDr2mXdnpd7Pot
7Z3V5p9wCl/dTL9WK6bcxWsccp4vO1/Cf3JrMzqJEk+NMVZR5msfyisdN0bXb6wkuLHVLuK6sL63
e5Ko/wDpmpreMwkTgkQurmct1AOw+EcVUjnP/ON9zps+jzWbnT4rqH62jWmrIkV1YWvpKk0xjHpv
BbRn1UZhxUO0g+0cVXahL+Teo69rs+syXGsLrk06yWqw3/p27WFtYWksSWsZ5vMyypIk8UPLgW+L
itSqm0/l/wDIvy3BZWBhj06C8tpr2xkhkvVX0oo4LmSdbiNv3cnDTYpVbmHJj5L8XKqqEs9X/I2C
2F0bOewa3nSFLWS21ATqbadIoWFugd1Xn5ejpVQR6YVwGYqyqGtpfyC0zWtN1nTbS4k1G1WzhsLm
FNTMKRXcFpZROGlK2zcba7t/U3LgFajnTFUPdf8AQtU2hJb3as2jRSwoYJV1f0bOSD91F6qMKWfP
1dy4T1aljy64q9QPnnyfFpNvqt1q9tp+n3MkkEMuoOLEmWF2jliKXPouskbxsrIwBBG4xVD3/wCY
3k+w1K8027vXju7DiLhRbXLJ6jxpMkMcqxGOWZo5FZYo2ZzXZcVSbVfzx/LzT9M1K+W7nvP0THFL
e20FvKsqCaSOML+/WFPUVp05xluS9CAdsVRVt+cPkSaF2a6ninS4urU2RtLmS6MllIkc3GCGOR2A
9VTUDoTWhVwqqZ6p+YHlHS/T+uX/ABM8MVxbiOGaYyxzpNJG0QiRzJVLWRqLUigr9paqpZp35xfl
9fx3E0WpcLa3lEP1iWOREc1tEYiq8k4S6lDG4kClWJqKKTirJNC1/S9dsfrumyO8AcxuJYZbeVXW
hKvDOkcqGhBHJRUEEbHFUwxV2KuxV2KuxVJ/Nv8Ahr9CN/iTh+ifrFp6nq8uHrfWovq3Lh2+sen1
+H+b4a4q8w0//oW+uhfUKV+uRfof0f0nT61wsPT58f8Aozr6u3Lr8XLFV/lP/oW3h5j/AMO/Vqfo
Y/p/j9d5fon0V58fU+KnDj6npfHypy+PFUND/wBC2/UxT1vqHp3/AC9X9M/V+fo3P1z1PU/d/WvR
9anP99xpw/YxVM9K/wCVBf49sP0b/wApR6lz+jeH6S9Dn9XP1n0a/wCiU9KvKm1f8rFWNah/0Lt+
g3/QXqfWv0Xdfof0/wBKfV+HoXnX1v8ARf8Alq/vtv7yv7WKor8nf+VQ/p+y9f0v+Vpf7kfrfq/X
vW5/Wrj1/wDev4ufp8uPr/6R6VOffFXoFh/yqL9Df7hf0T+if0lY8/0P6XpfpD6zF9S9T6l+36/p
05bePw4q8r85/wDKnvQP+BPR/wAafUtP+q0/Svqfo/0Y/Q4/V/8ASa/VuH9x+8pT1P3fPFUy0D/o
XD9AfB6/pfUbb67z/SlfT+o3FP7v91X6v9a9T0tv7yvfFWRTf8qA/SHl/n9W+ufpu7/QPH61X9Lf
WV+s/Z2/v+FPU/d/Z47UxVd52/5UT6Wt/wCJOFP0lH+mvQ+u8/r/AOj248vqnx0+pc+XH4Ptcvj5
Yqlup/8AQudPMH1/+5pefp2v6T9D1Pr0H1v7P7v6x9a+r19P95x4U+DjiqfQf8qt/wCVf6V9e9f/
AAr6l7+h/wBIfWv7n6tec+Ffj+r/AFD1vR5belxpvxxVJdD/AOhdf8GP+ivT/wAMfWbvnT9IU+sf
oqT61Xn+9r+j/Ur+Hx4qk+rf9CrfUJ/0pT6n9Yn/AL79Lf39LX1vq9d/sfV/7n/dfT4OWKsq80f8
qN9fQ/0z6fH9GH9Deh9d+rfoz6tL9r6t+49P6t63H1P2eVO+KoK9/wCVCfUIPr3rV9K59H1v0t+k
P+OtF6/2v9M9b9JcKV/eV5cfh5YqoH/oXj/Dq8uX6I4Lx5/pTl6dNK4fa/e8eP6O4+1Kbc8VYN5j
/wCVP/pWw/Rvq/4L9eL/ABb6v6Y9T1P3P1b1fU/3X/d8vrPw8a+j+854q9X0v/lRX+AdGp+h/wDB
vOX9C/pbh6Xqeo/q8P0h8fPnzrX4sVSWf/lXX+OfOn+JvrH1n63B9X/3v9Cn6KsqfVvQ/c/W/wCT
0/8ASKfZ2xVDXf8A0Lh6d99er6X1WP6/9Z/SvH0frNtXl6m3q/WfQ9f/AHZy/vf2sVa8wf8AQtH6
Pvv036PpfD9d9X9IfWeX1+660/0j1Prf1jnT4qfa+DjirI/NP/KlvqWkfpr6v9W/RT/oT0PrFf0b
W3/uPqvxdfS9Lj8fXh+1irFIv+hZf0Y3H1P0d9duvrXP9M+n9Y9Sz+tfXOW3per9U5ev+65ce9cV
Z9+U3/KtP8Kf8g79P9A/WZPU9P1+X1mi8/U+s/vuXHj9v9mlNqYqzPFX/9k=
51.000000
66.000000
Picas
1
False
False
Helvetica
Helvetica
Medium
Open Type
001.006
False
Helvetica.otf
Black
Default Swatch Group
0
application/postscript
endstream
endobj
11 0 obj
<< /S /URI /URI (www.pnas.org/cgi/doi/10.1073/pnas.1319538111)
>>
endobj
12 0 obj
<< /Filter /FlateDecode /Subtype /Type1C /Length 2 0 R
>>
stream
htW \fb8h .Xq¦HEފD
R֪(V@Ѡ\K]n)FclnQ\{{o9FN0wTQ#N[w(-C:>K~}(ukԊvSZ>gn>\m=$@9+#%M0[(5#+3#K6kN`B.+-;-+_8t:.'K?+m>k.c.>}QFN^f.2V_4k`F.f/INJOS_`KўQx)
Ib#cInJJ=pSGr^3Fd2v*;(Kmiփe9ύಸ*·;ݻ۞m?cIסC'9sk ^_yRuW
*Uw60Ju
1Z㮣K~ p< D+<? ®|"(`%= @J~
wb^1^*>,_-M
%BW͓Iu$8
xeI(.=j2w=
]dd#ok_|4p3Y↭B?wTs*\}M@}L