Similar title to Le Galliard, J.-F., Ferriere, R., & Dieckmann, U. ORCID: https://orcid.org/0000-0001-7089-0393 (2003). The adaptive dynamics of altruism in spatially heterogeneous populations. Evolution 57 (1) 1-17. 10.1111/j.0014-3820.2003.tb00211.x.