Similar title to Gyllenberg, M., Metz, J.A.J., & Service, R. (2011). When do optimisation arguments make evolutionary sense? In: The Mathematics of Darwin's Legacy. Eds. Chalub, FACC & Rodrigues, JF, Basel: Springer. ISBN 978-3-0348-0121-8 10.1007/978-3-0348-0122-5_12.